Homo floresiensis was recovered from Late Pleistocene depositson the island of Flores in eastern Indonesia, but has the stature,limb proportions and endocranial volume of African PlioceneAustralopithecus [1]. The holotype of the species (LB1), excavated in2003 from Liang Bua, consisted of a partial skeleton minus thearms. Here we describe additional H. floresiensis remains excavatedfrom the cave in 2004. These include arm bones belonging tothe holotype skeleton, a second adult mandible, and postcranialmaterial from other individuals. We can now reconstruct the bodyproportions of H. floresiensis with some certainty. The findsfurther demonstrate that LB1 is not just an aberrant or pathologicalindividual, but is representative of a long-term populationthat was present during the interval 95–74 to 12 thousand yearsago. The excavation also yielded more evidence for the depositionalhistory of the cave and for the behavioural capabilities ofH. floresiensis, including the butchery of Stegodon and use of fire.

Bones of the lower extremity have been recovered for up to nine different individuals of 'Homo floresiensis' - LB1, LB4, LB6, LB8, LB9, LB10, LB11, LB13, and LB14. LB1 is represented by a bony pelvis (damaged but now repaired), femora, tibiae, fibulae, patellae, and numerous foot bones. LB4/2 is an immature right tibia lacking epiphyses. LB6 includes a fragmentary metatarsal and two pedal phalanges. LB8 is a nearly complete right tibia (shorter than that of LB1). LB9 is a fragment of a hominin femoral diaphysis. LB10 is a proximal hallucal phalanx. LB11 includes pelvic fragments and a fragmentary metatarsal. LB13 is a patellar fragment, and LB14 is a fragment of an acetabulum. All skeletal remains recovered from Liang Bua were extremely fragile, and some were badly damaged when they were removed temporarily from Jakarta. At present, virtually all fossil materials have been returned, stabilized, and hardened. These skeletal remains are described and illustrated photographically. The lower limb skeleton exhibits a uniquely mosaic pattern, with many primitive-like morphologies; we have been unable to find this combination of ancient and derived (more human-like) features in either healthy or pathological modern humans, regardless of body size. Bilateral asymmetries are slight in the postcranium, and muscle markings are clearly delineated on all bones. The long bones are robust, and the thickness of their cortices is well within the ranges seen in healthy modern humans. LB1 is most probably a female based on the shape of her greater sciatic notch, and the marked degree of lateral iliac flaring recalls that seen in australopithecines such as "Lucy" (AL 288-1). The metatarsus has a human-like robusticity formula, but the proximal pedal phalanges are relatively long and robust (and slightly curved). The hallux is fully adducted, but we suspect that a medial longitudinal arch was absent.

'Homo floresiensis' is an endemic hominin species that occupied Liang Bua, a limestone cave on Flores in eastern Indonesia, during the Late Pleistocene epoch. The skeleton of the type specimen (LB1) of 'H. floresiensis' includes a relatively complete left foot and parts of the right foot. These feet provide insights into the evolution of bipedalism and, together with the rest of the skeleton, have implications for hominin dispersal events into Asia. Here we show that LB1's foot is exceptionally long relative to the femur and tibia, proportions never before documented in hominins but seen in some African apes. Although the metatarsal robusticity sequence is human-like and the hallux is fully adducted, other intrinsic proportions and pedal features are more ape-like. The postcranial anatomy of 'H.  floresiensis' is that of a biped, but the unique lower-limb proportions and surprising combination of derived and primitive pedal morphologies suggest kinematic and biomechanical differences from modern human gait. Therefore, LB1 offers the most complete glimpse of a bipedal hominin foot that lacks the full suite of derived features characteristic of modern humans and whose mosaic design may be primitive for the genus 'Homo'. These new findings raise the possibility that the ancestor of 'H.  floresiensi's was not 'Homo erectus' but instead some other, more primitive, hominin whose dispersal into southeast Asia is still undocumented.

Evidence from Liang Bua, a limestone cave on the island of Flores in East Indonesia, provides a unique opportunity to explore the long term relationship between hominins and their environment. Occupation deposits at the site span ῀95 ka and contain abundant stone artefacts, well preserved faunal remains and evidence for an endemic species of hominin: 'Homo floresiensis'. Work at the site included detailed geomorphological and environmental analysis, which has enabled comparisons to be drawn between changes in the occupational intensity in the cave, using stone tool and faunal counts, and changes in the environmental conditions, using the characteristics of the sedimentary layers in the cave and speleothem records. These comparisons demonstrate that 'H. floresiensis' endured rapidly fluctuating environmental conditions over the last ῀100 ka, which influenced the geomorphological processes in the cave and their occupational conditions. The intensity of occupation in the cave changed significantly between 95 and 17 ka, with peaks in occupation occurring at 100–95, 74–61 and 18–17 ka. These correlate with episodes of channel formation and erosion in the cave, which in turn correspond with high rainfall, thick soils and high bio-productivity outside. In contrast, periods of low occupational intensity correlate with reduced channel activity and pooling associated with drier periods from 94 to 75 and 36 to 19 ka. This apparent link between intensity of hominin use of the cave and the general conditions outside relates to the expansion and contraction of the rainforest and the ability of 'H. floresiensis' to adapt to habitat changes. This interpretation implies that these diminutive hominins were able to survive abrupt and prolonged environmental changes by changing their favoured occupation sites. These data provide the basis for a model of human–environment interactions on the island of Flores. With the addition of extra data from other sites on Flores, this model will provide a greater understanding of 'H. floresiensis' as a unique human species.

The announcement of a new species, 'Homo floresiensis', a primitive hominin that survived until relatively recent times is an enormous challenge to paradigms of human evolution. Until this announcement, the dominant paradigm stipulated that: 1) only more derived hominins had emerged from Africa, and 2) 'H. sapiens' was the only hominin since the demise of 'Homo erectus' and 'Homo neanderthalensis'. Resistance to 'H. floresiensis' has been intense, and debate centers on two sets of competing hypotheses: 1) that it is a primitive hominin, and 2) that it is a modern human, either a pygmoid form or a pathological individual. Despite a range of analytical techniques having been applied to the question, no resolution has been reached. Here, we use cladistic analysis, a tool that has not, until now, been applied to the problem, to establish the phylogenetic position of the species. Our results produce two equally parsimonious phylogenetic trees. The first suggests that 'H. floresiensis' is an early hominin that emerged after 'Homo rudolfensis' (1.86 Ma) but before 'H. habilis' (1.66 Ma, or after 1.9 Ma if the earlier chronology for 'H. habilis' is retained). The second tree indicates 'H. floresiensis' branched after 'Homo habilis'.

This study examines trends in stone tool reduction technology at Liang Bua, Flores, Indonesia, where excavations have revealed a stratified artifact sequence spanning 95 k.yr. The reduction sequence practiced throughout the Pleistocene was straightforward and unchanging. Large flakes were produced off-site and carried into the cave where they were reduced centripetally and bifacially by four techniques: freehand, burination, truncation, and bipolar. The locus of technological complexity at Liang Bua was not in knapping products, but in the way techniques were integrated. This reduction sequence persisted across the Pleistocene/Holocene boundary with a minor shift favoring unifacial flaking after 11 ka. Other stone-related changes occurred at the same time, including the first appearance of edge-glossed flakes, a change in raw material selection, and more frequent fire-induced damage to stone artifacts. Later in the Holocene, technological complexity was generated by "adding-on" rectangular-sectioned stone adzes to the reduction sequence. The Pleistocene pattern is directly associated with 'Homo floresiensis' skeletal remains and the Holocene changes correlate with the appearance of 'Homo sapiens'. The one reduction sequence continues across this hominin replacement.

Excavations at Liang Bua, Flores, Indonesia, have yielded evidence for an endemic human species, 'Homo floresiensis', a population that occupied the cave between ~95-17 ka. This discovery has major implications for early hominin evolution and dispersal in Africa and Asia, attracting worldwide interest. This preface describes the rationale for the excavations in historical, geographical, and wider research contexts, as well as the methods used. It also introduces the other papers on aspects of Liang Bua research that feature in this edition of the 'Journal of Human Evolution'.

Morwood et al discuss the discovery of the fossilised skeletal remains of what scientists nicknamed as the hobbits. Scientists believe that the skull represents a new human species called the Homo floresiensis, which were found in a cave on Flores. The species existed alongside modern humans as recently as 13,000 years ago, yet may descend from Homo erectus, which arose some two million years ago.

Song Gupuh, a partially collapsed cave in the Gunung Sewu Limestones of East Java, Indonesia, contains over 16 m of deposits with a faunal sequence spanning some 70 ka. Major changes in the range of animals represented show the impact of climate change and humans. The Terminal Pleistocene and Early Holocene was a period of maximum biodiversity. Human use of Song Gupuh and other cave sites in the region also intensified significantly from ca. 12 ka, together with a new focus on exploitation of small-bodied species (macaque monkeys and molluscs), the first evidence for import of resources from the coast, and use of bone and shell tools. Human activity, especially after the onset of the Neolithic around 2.6 ka, subsequently contributed to a progressive loss of many species from the area, including tapir, elephant, Malayan bear, rhino and tiger, and this extinction process is continuing. We conclude by discussing the biogeographical significance of Song Gupuh in the context of other sites in Java (e.g. Punung, Wajak) and further afield (e.g. Liang Bua).

Several bones of the upper extremity were recovered during excavations of Late Pleistocene deposits at Liang Bua, Flores, and these have been attributed to 'Homo floresiensis'. At present, these upper limb remains have been assigned to six different individuals - LB1, LB2, LB3, LB4, LB5, and LB6. Several of these bones are complete or nearly so, but some are quite fragmentary. All skeletal remains recovered from Liang Bua were extremely fragile, but have now been stabilized and hardened in the laboratory in Jakarta. They are now curated in museum-quality containers at the National Research and Development Centre for Archaeology in Jakarta, Indonesia. These skeletal remains are described and illustrated photographically. The upper limb presents a unique mosaic of derived (human-like) and primitive morphologies, the combination of which is never found in either healthy or pathological modern humans.